Porth's Essentials of Pathophysiology, 4e

831

Organization and Control of Neural Function

C h a p t e r 3 4

POSTERIOR Dorsal view

Future neural crest

Neural plate Somites

ANTERIOR

1

Neural plate

Ectoderm

Neural folds

Notochord

Endoderm

Rostral neuropore

Mesoderm

Neural groove

Neural folds

1

Cut edge of amnion

Neural crest

Ectoderm

2 3

2

Neural tube Yolk sac wall Neural groove Neural folds

Somites (mesoderm)

Mesoderm Endoderm

Notochord

Neural groove

Caudal neuropore

Neural tube

Neural crest

Mesoderm

Neural plate

Ectoderm

3

POSTERIOR Dorsal view

A

B

Notochord

Endoderm

Transverse sections

FIGURE 34-5. Folding of the neural tube. (A) Dorsal view of a six-somite embryo (22 to 23 days) showing the neural folds, neural groove, and fused neural tube.The anterior neuropore closes at about day 25 and the posterior neuropore at about day 27. (B) Three cross-sections taken at the levels indicated in A.The sections indicate where the neural tube is just beginning to form.

Segmental Organization Developmentally, the basic organizational pattern of the nervous system is that of a longitudinal series of segments, each repeating the same fundamental pattern that is retained in postnatal life. The CNS and its associated peripheral nerves consist of approximately 43 segments,

CSF-filled cavities, the first and second (lateral) ven- tricles. The remaining diencephalic portion of the neu- ral tube develops into the thalamus and hypothalamus. The neurohypophysis (posterior pituitary) grows as a midline ventral outgrowth at the junctions of segments 1 and 2. A dorsal outgrowth, the pineal body, develops between segments 2 and 3.

Mesencephalon (Midbrain)

Prosencephalon (Forebrain) Optic vesicle

Rhombencephalon (Hindbrain)

(developing eye)

Prosencephalon (Forebrain)

Developing heart

Mesencephalon (Midbrain)

Rhombencephalon (Hindbrain)

FIGURE 34-6. Frontal and lateral views of a 5-week-old embryo showing the brain vesicles and three embryonic divisions of the brain and brain stem.

Spinal cord

Frontal view

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